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Recent works in multi-agent systems have identified agent behaviors that can develop and sustain mutually beneficial cooperative relationships with like-minded agents and can resist exploitation from selfish agents. Researchers have proposed the use of a probabilistic reciprocity scheme that uses summary information from past interactions to decide whether or not to honor a request for help from another agent. This behavior has been found to be close to optimal in homogeneous groups and outperform exploiters in mixed groups. A major shortcoming of these experiments, however, is that the composition of the group in term of agent behaviors is fixed. We believe that real-life rational agents, on the contrary, will change their behaviors based on observed performances of different behavioral traits with the goal of maximizing performance. In this paper, we present results from experiments on two distinct domains with population groups whose behavioral composition changes based on the performance of the agents. Based on the experimental results, we identify ecological niches for variants of exploitative selfish agents and robust reciprocative agents.